Find the Predecessor: Identifying the Number Before 40,300

To solve the problem of finding what number comes before 40,300, we need to find the predecessor of 40,300. This can be done by subtracting 1 from 40,300.

Let's perform this calculation:

• Given number: $40,300$
• Predecessor: $40,300 - 1$
• Calculating: $40,300 - 1 = 40,299$

Therefore, the number that comes before 40,300 is $40,299$.

Looking at the provided answer choices, choice #4 offers the number 40,299, which matches our calculation. Thus, the correct answer is $40,299$.